CLASSE: GASTROPODA: TERRESTRE
FAMÍLIA: SIMPULOPSIDAE
ESPÉCIE: Leiostracus subtuszonatus (Pilsbry, 1899)
Tamanho médio: 25 mm
Habitat: Mata Atlântica, vive em árvores e arbustos
Ocorrência: Brasil (Bahia, Minas Gerais)
Localidade tipo: NEOTYPE: Ilhéus , Bahia, ~52 m de altitude (Salvador & Cavallari, 2013)
Nome original: Drymaeus (Leiostracus) onager var. subtuszonata Pilsbry, 1899
Etimologia: o nome da espécie “subtuszonatus” indica que esse é o padrão de cores “típico” das bandas axiais subitamente interrompido por uma faixa em espiral, uma útil característica diagnóstica desta espécie
Redescrição:
(Salvador & Cavallari, 2013): “Shell medium-sized, sub-conical; width ~1/2 shell length. Spire top colour (~3 whorls, including protoconch) whitish to yellowish brow; remaining whorls whitish with thick brown axial bands, sometimes forked. On body whorl, these axial bands are interrupted by a broad yellow or whitish spiral band flanked by brown bands (the upper brown band, usually much narrower, may not be present on some specimens).
Colour pattern visible from the inside of shell; peristome and umbilical region white to light yellowish brown. Spire angle ~45°. Protoconch (~1½ whorls) sculptured by fine parallel axial wrinkles on the upper portion of whorls, but this sculpture stops in the middle portion and gives place to very fine parallel spiral lines; transition to teleoconch unclear. Teleoconch smooth, except for growth lines. Whorls profile convex. Suture well-marked, slightly oblique (diagonal) to columellar axis. Aperture large, oval, slightly prosocline (~15º with columellar axis); ~2/5 shell length, ~2/3 shell width. Peristome reflected, especially on columellar region, partially covering umbilicus. Body whorl ~3/5 shell length. Umbilicus rimate.”
Fotos:
(1-4) Natural History Museum, London: prováveis PARALECTOTYPES de Drymaeus (Leiostracus) onager subtuszonatus Pilsbry, 1899, NHMUK 20130094, coll. Hugh Cuming



Figura 1: Salvador & Cavallari, 2013: Leiostracus subtuszonatus: (14–17) NEOTYPE MZSP 108040, 28.5 mm; (18) SEM imagem da protoconcha MNHN IM-2012–2007; (19) MZSP 27650, 21.5 mm; (200 juvenil, MZSP 110707, 24 mm; (21) SMF 284019, 28 mm; (22) SMF 284019, 25.5 mm; (23) MNHN IM-2012–2007, 25.5 mm; (24) MNHN IM-2012–2007, 28 mm

Figura 2: Salvador, Rodrigo & Cavallari, Daniel & Breure, Bram. (2014): (1–2) Leiostracus subtuszonatus ANSP 25963; (H = 25 mm); (3–4) Leiostracus subtuszonatus NHMUK 20130094; (H= 27.5 mm); ( 5–6) L. subtuszonatus NHMUK 20130094, espécime juvenil, (H=24.5 mm); ( 7–8) Leiostracus subtuszonatus NHMUK 20130094; (H=29 mm); (9) Reprodução da figura de Leiostracus subtuszonatus de Reeve (1848: pl. 45, fig. 284), designada como LECTOTYPE; (10) Reprodução da figura de L. subtuszonatus de Pilsbry (1899: 95, pl. 14, fig. 17)

Figura 3: Salvador, R. B. & Cavallari, D. C. (2014): (9) Leiostracus onager (Beck, 1837), NEOTYPE (MZSP 29794; H = 25.5 mm, D = 12 mm); (10) Leiostracus subtuszonatus (Pilsbry, 1899) (MZSP 108040; H = 28.5 mm, D = 13 mm); (11) Leiostracus clouei (Pfeiffer, 1856), LECTOTYPE (NHMUK 1975491; H = 22 mm, D = 11 mm)
Fontes:
Descrição original: Pilsbry, H.A. 1899. Manual of Conchology. Second Series: Pulmonata. Vol. 12. American Bulimulidae: North American and Antillean Drymaeus, Leiostracus, Orthalicinae and Amphibuliminae. Academy of Natural of Sciences of Philadelphia, Philadelphia. 258 pp
Simone, L. R. L. (2006). Land and Freshwater Molluscs of Brazil. Editora Gráfica Bernardi, FAPESP. São Paulo, 390 pp. página(s): 122, Fig. 387
Salvador, Rodrigo & Cavallari, Daniel. (2013). Taxonomic revision of Leiostracus onager and Leiostracus subtuszonatus (Gastropoda: Pulmonata: Orthalicidae). Journal of Conchology. 41. 511-518. https://www.researchgate.net/publication/259150873_Taxonomic_revision_of_Leiostracus_onager_and_Leiostracus_subtuszonatus_Gastropoda_Pulmonata_Orthalicidae
Salvador, Rodrigo & Cavallari, Daniel & Breure, Bram. (2014). Corrigendum to “Taxonomic Revision of Leiostracus Onager and Leiostracus subtuszonatus (Gastropoda: Pulmonata: Orthalicidae)” by Salvador & Cavallari (2013). Journal of Conchology. 41. 627-628
Salvador RB, Simone LRL (2021) New records of Orthalicoidea land snail species for the state of Minas Gerais, southeastern Brazil (Gastropoda, Stylommatophora). Check List 17(2): página(s) 395-399, disponível online em https://doi.org/10.15560/17.2.395
MolluscaBase: https://www.molluscabase.org/aphia.php?p=taxdetails&id=1292075